The winners of this year's British Wildlife Photography Awards have been announced. The competition's 12 categories cover everything from marine life and animal behaviour to creepy crawlies and urban wildlife, revealing the surprising diversity of Britain's wildlife.

This year's winner is Lee Acaster from Suffolk with his photo of a greylag goose by the Thames. Acaster said: "I was set up for shooting a stormy cityscape with a manual focus wide angle lens on when I came across the goose sat on the river wall. Expecting it to fly away as I got nearer, I was surprised to find that it was very happy to stay where it was, even when I got very close.

"It was technically incredibly difficult to get the shot, holding a flash out in one hand and my camera in the other, trying to focus on the goose by moving closer to him without scaring him away. I ended up being just a few inches away from him for the final image. He was still happily stood on the wall as I left, probably wondering what on earth the strange man with the flashing light had been doing."

British Wildlife Photography Awards
Overall Winner: The Tourist by Lee AcasterLee Acaster

The winning photos and many more are published in the British Wildlife Photography Awards Collection 4 (AA Publishing £25) and can be seen in a free exhibition at the Mall Galleries in London from 2 September.

IBTimesUK presents a gallery of the winning images in each category.

British Wildlife Photography Awards
Habitat Winner: A Life at Sea - Nesting Gannets, Shetland Isles by Ruth AsherRuth Asher
British Wildlife Photography Awards
Seasons Winner: Spring is in the Air by Andrew ParkinsonAndrew Parkinson
British Wildlife Photography Awards
Behaviour Winner: Otter and Puffin by Richard ShucksmithRichard Shucksmith
British Wildlife Photography Awards
Wild Woods Winner: Autumn Jewels by Peter CairnsPeter Cairns
British Wildlife Photography Awards
Coast and Marine Winner: Big Blues by Alexander MustardAlexander Mustard
British Wildlife Photography Awards
Hidden Britain Winner: Window Gnat by Susie HewittSusie Hewitt
British Wildlife Photography Awards
Documentary Winner: Hazel Dormouse Monitoring by Nick UptonNick Upton
British Wildlife Photography Awards
Portraits Winner: Shag resting - Farne Islands, Northumberland by Steven FairbrotherSteven Fairbrother
British Wildlife Photography Awards
Portraits Highly Commended: Waving Seal by Luke WilkinsonLuke Wilkinson
British Wildlife Photography Awards
Portraits Highly Commended: Badger - Brock over Brook by Russell SavoryRussell Savory
British Wildlife Photography Awards
Portraits Commended: Owl by Matt BinsteadMatt Binstead
British Wildlife Photography Awards
Black & White Winner: Blue on Black, blue shark by Alexander MustardAlexander Mustard
British Wildlife Photography Awards
Black & White Commended: Swan in Morning Mist by Andrew KaplanAndrew Kaplan
British Wildlife Photography Awards
Close to Nature Winner: Grey Seal Claws by Jim GreenfieldJim Greenfield
British Wildlife Photography Awards
Botanical Winner: Green and red telephone box by Philip BraudePhilip Braude
British Wildlife Photography Awards
Under 12 Winner: Dipper with Grubs by William Bowcutt, 11William Bowcutt
British Wildlife Photography Awards
Age 12 to 18 Winner: On the Prowl by Joshua Burch, 16Joshua Burch